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4.4 Meiosis: Cell Divisions That Halve Chromosome Number   105









                                                                                               Figure 4.15  To aid visualization
                                                                                               of the chromosomes, the figure is
                                                                                               simplified in two ways: (1) The nuclear
                                                                                               envelope is not shown during prophase
                                                                                               of either meiotic division. (2) The
                                                                                               chromosomes are shown as fully
                                                                                               condensed at zygotene; in reality, full
                                                                                               condensation is not achieved until
                                                                                               diakinesis.


                                  Prophase I: Diplotene         Prophase I: Diakinesis
                                  1.  Synaptonemal complex dissolves.  1.  Chromatids thicken and shorten.
                                  2. A tetrad of four chromatids is visible.  2. At the end of prophase I, the nuclear
                                  3. Crossover points appear as chiasmata,      membrane (not shown earlier) breaks
                                      holding nonsister chromatids together.      down, and the spindle begins to form.
                                  4. Meiotic arrest occurs at this time in many
                                      species.


















                                   Telophase I                  Interkinesis
                                   1.  The nuclear envelope re-forms.  1.  This is similar to interphase with one
                                   2. Resultant cells have half the number of      important exception: No chromosomal
                                       chromosomes, each consisting of two      duplication takes place.
                                       sister chromatids.       2. In some species, the chromosomes
                                   3. Cytokinesis separates the daughter      decondense; in others, they do not.
                                       cells (not shown).



















                                  Telophase II                  Cytokinesis
                                  1.  Chromosomes begin to uncoil.  1. The cytoplasm divides, forming four
                                  2. Nuclear envelopes and nucleoli (not      new haploid cells.
                                      shown) re-form.
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